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Yet, little is known about either the genetic origins or the lifeways of these border peoples. Han/non-Han interactions were engrained among the border regions of ancient Imperial China.

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10Institute of Artificial Intelligence, Xiamen University, Xiamen, China.9State Key Laboratory of Marine Environmental Science, Xiamen University, Xiamen, China.8State Key Laboratory of Cellular Stress Biology, School of Life Sciences, Xiamen University, Xiamen, China.7Xinjiang Institute of Cultural Relics and Archaeology, Urumqi, China.6Department of Archaeology and Anthropology, University of Chinese Academy of Sciences, Beijing, China.5Ministry of Education Key Laboratory of Contemporary Anthropology, Department of Anthropology and Human Genetics, School of Life Sciences, Fudan University, Shanghai, China.4Center for the Belt and Road Archaeology and Ancient Civilizations, Fudan University, Shanghai, China.3Institute of Archaeological Science, Fudan University, Shanghai, China.2Department of Anthropology and Ethnology, School of Sociology and Anthropology, Institute of Anthropology, Xiamen University, Xiamen, China.1MOE Laboratory for National Development and Intelligent Governance, Department of Cultural Heritage and Museology, Fudan University, Shanghai, China.Edward Allen 1† Yao Yu 1† Xiaomin Yang 2† Yiran Xu 3,4 Panxin Du 5 Jianxue Xiong 5 Dian Chen 6 Xiaohong Tian 7 Yong Wu 7 Xiaoli Qin 1 Pengfei Sheng 1,3,4* Chuan-Chao Wang 2,8,9,10* Shaoqing Wen 1,3,4*














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